Led by Gendo Ikari, the forces of the apocalyptic death cult NERV clash with WILLE, a resistance organisation comprises of former NERV employees and led by Shinji`s guardian, Katsuragi Misato. As WILLE tries to revert the red taint that contaminates Paris, a pair of Evangelions piloted by Asuka and Mari are all that stand against NERV`s own horde of Evas.